Miles Axlerod is an ex oil baron who has sold off his fortune, converted himself into an electric vehicle, and has devoted his life to finding the renewable, clean-burning energy source of the future—ultimately discovering what he believes is the fuel everyone should be using. Axlerod is also the car behind the World Grand Prix, a three-country race he created that attracts the world’s top athletes, but it's really an excuse to promote his new clean burning wonder-fuel, Allinol. During most of the movie, Professor Z seems to be the main antagonist, but it turns out that Miles Axlerod was the main antagonist, and Professor Z was the secondary antagonist.
- Miles Axlerod was voiced by Eddie Izzard, who also voiced Lord Voldemort (The LEGO Batman Movie.
- He will encounter Winnie the Pooh, Thomas, Twilight Sparkle, Ash Ketchum, Littlefoot and their friends in Pooh's Adventures of Cars 2, where at the end, he's arrested.
- Miles Axlerod will encounter Ryan, Crash, Thomas and their friends in Ryan's, Thomas' and Crash's Adventures of Cars 2, where at the end, he was arrested.